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Mares are female horses.  The mare at the front and center of this picture is a band mare.  She eventually went off to a different stallion and became a lead mare.  A lead mare is the mare that all other mares of a band look up to.  Lead mares are responsible for taking the horses to food, water, and safe places.  The paths that lead mares take eventually become horse trails.

This is the other mare in the picture at left. She was the lead mare of this band.